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when the first transistor turns on thru the slightest imbalance, the voltage at the collector starts to be pulled down and this is felt through the capacitor at the base of the other transistor as a negative voltage - thus turning that transistor off its collector voltage starts to increase towards the su voltage turning the first transistor more fully on. When the first transistor is fully turned on, and the collector voltage is no longer going negative, the capacitor now starts to charge up thru its base resistor. When the voltage gets high enough the second transistor starts turn on, it's collector then starts to go less positive, this is felt thru a capacitor as a negative change on the base of the first transistor, thus starting to turn it off, and the cycle continues.
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